The Hyphae Patrol quest from Marasmius doesn’t work if the Shakin’ Shrooms world quest is active.
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There is a bug that the quest will not track unless you exit the vehicle or exit the dialogue before entering the vehicle. Was it just not tracking your enemy roots? It may be unrelated to the world quest, if so.
The counter started working after I completed the WQ.
Ah ok. It could also have worked because you had to initiate the dialogue with Marasmius and the “talk to Marasmius” option was finally completed then.